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Is it possible for another player(player 2) to join a nearly complete game of borderlands 2 to (for example) defeat a boss?

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Yes, they can, although weapon damage ramps up at an exponential rate, so unless player 2 is close to or above the same level as player 1, they won't be much help against the boss.

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If he stays (relatively) out of the fight though, wouldn't P2 get the mountain of experience that comes with defeating higher level enemies? – Robotnik Apr 30 '13 at 3:24
@Robotnik Yes, although it's not as much as you might think. A month or two ago, I, as a level 24, joined two level 50s in killing Vermivorous the Invincible a few times. After killing him 5 or six times, I think I was level 28. While I didn't completely stay out, I certainly wasn't helping much. My criticals were doing 48 damage. As Zero. – SaintWacko Apr 30 '13 at 3:30
If you're in it for the xp, the warrior goes a lot faster, he gives about the same amount of xp, but he dies a lot faster. Unless you use the infinity pistol glitch ofcourse. – Kevin Apr 30 '13 at 11:54

Yes, although if the second player has never played before, he will be level 1 and will most likely get killed very fast by those high level enemies.

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Yes, and you may want to see How does mission progress work in multiplayer? which discusses how plot/mission progression works. Also, How does experience work in co-op?

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