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Occasionally I'll face a Charizard or Venusaur in GBL that has dark striping like this:

enter image description here

How do they get those markings, or what is the significance of them?

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These are clone Pokémon, which were available in 4-star raids during Pokémon Day 2020. They are just cosmetic variants of Venusaur, Charizard, Blastoise, and Pikachu.

Clone Blastoise, Charizard, and Venusaur

The evolved clone starters came with their respective Community Day charge move, so you can expect them to have it even at low MMR.

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    I assume they are a reference to the first Pokemon movie?
    – Stef
    Feb 6 at 13:02
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    @Stef Definitely yes.
    – pinckerman
    Feb 6 at 13:31
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    Technically, it was to promote the 2019 Remake of the first movie ;) Feb 6 at 16:47

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